What does «substrato» mean in Spanish?

In Philosophy, essence, substance, permanent characteristics of a thing.
In geology, the ground beneath an overlying layer; subsoil.

Examples of use in Spanish: "el substrato de un manto".
Deep origin of something.

Examples of use in Spanish: "el substrato ibérico de la población".
Potting soil.

Examples of use in Spanish: "debes conseguir un buen sustrato para plantar estas plantas".
In ecology, part of the biotope where certain living beings carry out their vital functions.
Molecule on which an enzyme acts.
A chemical species that is explicitly considered to be the target of the action of other reagents.
In biology, a place used for planting plants.
In photography, a bath applied to the substrate that allows adhesion between the photosensitive layer and the glass or plastic materials.
In Language, a language that influences another language in the same territory.
In Architecture, material that remains under a superimposed layer that serves as a seat. Also called underlayment.